At 183.9 inches (4.67 m) in length, the FJ Cruiser is a mid-size SUV offered by Toyota in North America, slotting between the smaller RAV4 and larger Highlander. Toyota sources many components from other vehicles to produce the FJ including shared engines and transmissions found in the Tacoma, Tundra and 4Runner as well as suspension components used in the Prado, Hilux, Tacoma, and 4Runner.

The 4x4 FJ Cruiser has 9.6 inches (24 cm) of ground clearance, 34° approach and 30° departure angles, and a 27.4° breakover angle, with 8 inches (20 cm) of front and 9 inches (23 cm) of rear suspension travel. The FJ is also designed to drive through up to 27.5 inches (70 cm) deep of water and has a 5,000 lb (2,268 kg) towing capacity for both 4x4 and 4x2 variants. On road, the FJ accelerates to 60 mph (97 km/h) in 7.8 seconds and has 0.69 G of lateral grip tested with stock Bridgestone Dueler tires.
Structurally, the FJ Cruiser incorporates a body on frame truck style design similar, but not identical to that of the two-door Prado platform. The FJ Cruiser uses a high-mounted, double wishbone front suspension and stabilizer bar, and a 4-link rear suspension with lateral rod with coil springs and stabilizer bar. The 120-series Land Cruiser Prado, Tacoma, 4Runner, and Hilux share the same suspension parts as the FJ Cruiser. The FJ Cruiser uses power-assisted four-piston front and two-piston rear ventilated disc brakes with an anti-lock braking system (ABS), electronic brakeforce distribution (EBD), brake assist (BA) and is also equipped with vehicle stability control (VSC) and a specialized traction control system known as "A-TRAC". The A-TRAC system applies braking to control wheels that have lost traction, mimicking the performance of a locking differential without the "binding" that can make steering difficult in normal locking differential setups.

Toyota made revisions to the engine bay's inner fender aprons due to some bulging and cracking issues with the 2007 and some early 2008 models. The bulges and cracks in the inner fenders were attributed to a Toyota design flaw through certain vehicle identification numbers. Toyota addressed the problem and has changed the design.
The FJ Cruiser uses the 4.0-liter 1GR-FE DOHC V6. For the 2007 to 2009 model years, the engine came with single variable valve timing (VVT-i) which produces 239 hp (178 kW) at 5,200 RPM, and 278 lb⋅ft (377 N⋅m) of torque at 3,700 RPM on 91 octane fuel. For the 2010 model year, the FJ Cruiser's engine gained Dual VVT-i technology which adjusts timing on both intake and exhaust camshafts resulting in 259 hp (193 kW) at 5,600 RPM, and 270 lb⋅ft (366 N⋅m) on 87 octane, in addition to slightly improved fuel economy on 91 octane fuel. For model year 2011, these figures rose to 260 hp (194 kW) at 5600 RP…

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The FJ Cruiser was primarily developed for the North American market but has been sold in small numbers globally since its launch. On 25 November 2010, Toyota announced sales would begin in right-hand drive form for the Japanese market. Sales began on 4 December, replacing the outgoing Hilux Surf and selling alongside the Land Cruiser. Japanese-produced right-hand drive models have a different rear bumper to North American models with the license plate located on the bumper as opposed to under the tailgate door handle. In addition, right-hand drive versions have the side indicator markers located on the fender panels, rather than located on the side-view mirrors used for left-hand drive production FJ Cruisers. Sales of right-hand drive model began in Australia and New Zealand in 2011 available in only four-wheel drive with automatic transmission and a petrol engine.

Japanese market sales, never substantial to begin with, were halted in January 2018.

For North America, in 2007 Toyota Racing Development (TRD) produced 3,200 units as limited production TRD Special Edition FJ Cruisers. This model was released with the body painted in a black diamond pearl color, and with a matching black roof to set itself apart from its standard, white-roofed counterparts. The special edition model included TRD cat-back exhaust system, TRD Bilstein offroad tuned shocks, rock rails, 16-inch alloy gun-metal gray finished TRD wheels with BFGoodrich all-terrain tires, and TRD badges. The TRD Special Edition also included mechanical changes to synchronize the locking differential with the active traction control (resulting in the rear differential no longer overriding the active traction). Early production 2006 FJ Cruisers had an issue that caused the rear differential to override the active traction control, except in the TRD Special Editions. However, by November 2006, Toyota had changed production to incorporate synchronization of the locking differential with active traction control on all FJ Cruisers.

In 2009, subsequent to Toyota releasing the Trail Teams Special Edition FJ Cruiser for the production year of 2008, Toyota created the TRD package. This package was available on FJ Cruisers with the iceberg white monotone exterior color scheme, similar to the 2008 Trail Teams Special Editions, except without the addition of blacked-out trim pieces. Even though this package was known as the "TRD package", it did not include the same or as many features as the TRD Special Edition. Some of the features with this package were the TRD alloy wheels in a silver finish with the BFGoodrich all-terrain tires, as well as TRD logos and graphics on the exterior. Also, the TRD package was only offered on units with two-wheel drive for the 2008 and 2009 model years. Since 2011, Toyota offered the TRD package on the iceberg white FJ Cruisers but opened the package availability to both two- and four-wheel drive models. In 2009, 2,400 units were produced with the TRD package.
Beginning with the 2008 model year, Toyota manufactured 3,200 units of specially equipped FJ Cruisers designated as the Trail Teams Special Edition, which featured many of the same TRD options included on the TRD Special Edition, including the same 16-inch alloy wheels (but in a black finish as opposed to the gun-metal grey finish of the TRD model). The Trail Teams Special Edition was also equipped with BFGoodrich All-Terrain tires, as well as TRD/Trail Teams Bilstein shocks, and other similar options found on the TRD model. The number of Trail Teams Special Edition FJ Cruisers produced per year are as follows: 3,200 units for 2008 (as stated above); 1,500 for 2010; 2,500 in 2011; 2,500 in 2012; 2,500 in 2013; and 2,500 in 2014. No Trail Teams models were produced in 2009. However, Toyota released a TRD Package for that model year and it was used on the same Iceberg White painted FJ Cruisers that wer…

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